The upper molar cusps was relatively bunoselenomorph

The upper molar cusps was relatively bunoselenomorph

The upper molar cusps was relatively bunoselenomorph

He has got a short and you may straight postprotocrista labially situated. This new preprotocrista, seemingly straight, touches a well-developed parastyle. The latest metaconule is actually quite smaller that will be larger towards the M3. The new premetaconulecrista and also the fossae are simple in place of neocrista. The posterior fossa try angled, specifically into M3. Out of upright towards the M1, the newest postmetaconulecrista will get sigmoidal to the M3, bypassing the latest metacone. The fresh new labial cusps come into a comparable axis having seemingly straight cristae. The fresh paracone labial rib is actually outstanding, but versus anterior groove. There is absolutely no metacone labial rib. The brand new para poder-, meso-, and metastyles was outstanding. Brand new lingual cingulum is very weak to absent like the entostyle.

The fresh new registered build of your own fourth lower premolar is normal for derived base pecoran and you may crown pecoran when you look at the that have a well-put up mesolingual conid and you will posterior cristid , . Also, the clear presence of an enamel retailer into p1 excludes an attribution so you’re able to Prodremotherium or Mosaicomeryx . Even if the pearly whites are not because the bunoselenodont such as Amphitragulus elegans, the latest referred specimens have quite primitive have, particularly an incredibly quicker metastylid, a low crown, a small postentocristid, and even though laterally compacted, bulged lingual cuspids into all the way down molars. All these qualities exclude an association so you can both Dremotherium otherwise Bedenomeryx , . Also, the latest extremely small-size out of “Amphitragulus” quercyi is actually a pinpointing feature toward latest Oligocene ruminants away from Europe , , , .

Dremotherium cf. guthi from Gaimersheim1: step 1 remaining maxillary limbs to your tooth line M1-M3 (SNSB-BSPG 1952II4855). away from Mailhat: dos leftover P4-M2 (MNHN LIM154). Babameryx engesseri away from Gaimersheim1: step 3 kept fragmentary mandible that have m2 (SNSB-BSPG 1952II4818), (a) lingual, (b) labial, and you may (c) occlusal viewpoints. The scale bar try 2 cm.

Holotype referred specimens information

Holotype. Jehenne (1987) enjoys designated this new sample IPHEP LM.1969.MA.forty since the holotype. This can be a partial best maxillary bones that have P3-M3 (, Fig. 2), out-of La Milloque (MP29, France).

Called specimens. Mailhat MNHN LIM154 kept P4-M2 (this work, Fig. 6.2); Gaimersheim1 SNSB-BSPG 1952II4855 remaining maxillary that have M1-M3 (it really works, Fig. 6.1); SNSB-BSPG 1952II4870 proper m2.


All the way down long lasting dentition. The lower molar is extremely badly preserved. However, that it huge ruminant possesses a totally selenomorph trigonid. The newest cristid try elongated and cuspids is actually high towards the tooth a bit pleated. The new metastylid was ripped.

Higher long lasting dentition. New P4 was poorly maintained. Every lingual part are destroyed. The brand new anterolabial cone features a failing labial rib. The fresh new anterior stylid was busted. The latest posterior a person is small however, introduce. Zero neocrista obstructs the fossa. The fresh quadratic upper molars inside occlusal see incur a highly-put up metaconule a little smaller than the fresh protocone. New cusps have become selenomorph. The latest preprotocrista additionally the postmetaconulecrista is labially dependent and a little upright. This new postprotocrista try a lot of time, very rounded, and labially depending. It may be entirely remote (SNSB-BSPG 1952II4855) otherwise fused (MNHN LIM154) to the premetaconulecrista. The latter finishes at axis of mesostyle. Brand new posterior basin can also be possess a small metaconule flex to your M2, larger for the M3. Brand new paracone, possessing good paracone rib, while the metacone, labially apartment and you will concave, was aimed. The brand new mesostyle and you will parastyle are very well install. It form rounded small articles. The fresh new mesostyle try huge off M1 to M3. The brand new entostyle is poor or missing. There was neither an effective lingual cingulum close the latest protocone nor a beneficial metastyle building a highly-set up cuspid. However, short anterior and posterior cingula exist.


It ruminant is fairly complex in starting to be better-selenodont and achieving no lingual cingula during the higher molars, always within Lophiomeryx, Gelocus, Iberomeryx, Mosaicomeryx, and you may Babameryx , , , [61–63], . In comparison to Bachitherium, so it varieties lacks an anterior groove on the large labial paracone rib , , . New learnt fossils be similar to the complex stem Pecora and the brand new primitive crown Pecora (such as for example Amphitragulus, Dremotherium, Bedenomeryx). The brand new tooth is some pleated, while it is basically easy during the Prodremotherium. Additionally, the size of this type of teeth is a lot larger than in Prodremotherium regarding the dated Quercy series and you will Pech Desse . Tooth be selenomorph compared to “Amphitragulus” and you will Bedenomeryx , , as well as the parastyle is not that anteriorly projected than seen for the “Amphitragulus” feningrei , , . Looking at the neodiagnosis suggested because of the Mennecart , such fossils tend to be all of the demonstrating attributes of Dremotherium. The fresh new currently discussed species into the European countries try D. feignouxi (type of varieties) and D. guthi , , . The latter try a little smaller compared to the type variety , just like the fossils demonstrated right here. D. feignouxi only took place during the Agenian (MN1-MN2; ), whereas D. guthi lived into the most recent Oligocene (MP28-MP30; ). Yet not, within the absence of even more well-maintained matter, a definitive assignation towards the variety isn’t feasible.

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